Link to Google Scholar Profile


McClain, C.R. and Rex, M.A. (2001) The Relationship Between Dissolved Oxygen Concentration and Maximum Size in Deep-Sea Turrid Gastropods: An Application of Quantile Regression. Marine Biology, 139: 681-685

McClain, C.R., M. Fougerolle, M.A. Rex, and J. Welch. (2001) MOCNESS Estimates of the Size and Abundance of a Pelagic Gonostomatid Fish (Cyclothone pallida) off the Bahamas. Journal of the Marine Biological Association of the United Kingdom, 81:869-871


Linse, K., M. Schrödl, C.R. McClain, and L. Allcock (2003) Mollusca in the Antarctic deep sea-preliminary notes on their taxonomy, biogeography, and diversity. Reports on Polar and Marine Research (ANDEEP I & II: Antarctic benthic deep-sea biodiversity). 470:95-101


McClain, C.R., M.A. Rex, and N. Johnson (2004) Morphological Disparity as a Biodiversity Metric in Lower Bathyal and Abyssal Gastropod Assemblages. Evolution, 58:338-348

McClain, C.R. (2004) Connecting species richness, abundance, and body size in deep-sea gastropods. Global Ecology and Biogeography, 13: 327-334


McClain, C.R. and Etter, R.J. (2005) Mid-Domain Models As Predictors of Species Diversity Patterns: A Case Study Using Bathymetric Diversity Gradients in the Deep Sea. Oikos, 109:555-566

Rex, M.A., C.R. McClain, N.A. Johnson, R.J. Etter, J.A. Allen, P. Bouchet, A. Warén (2005) A Source-Sink Hypothesis for Abyssal Biodiversity. American Naturalist, 165:163-178 [Am. Nat. Highlight Paper of 2005]

McClain, C.R., M.A. Rex, and R. Jabbour. (2005) Deconstructing bathymetric patterns of body size in deep-sea gastropods. Marine Ecology Progress Series, 297:181-187

McClain, C.R. (2005) Bathymetric patterns of morphological disparity in deep-sea gastropods from the western North Atlantic Basin. Evolution, 59:1492-1499


McClain, C.R., A. Boyer, G. Rosenberg (2006) The island rule and the evolution of body size in the deep sea. Journal of Biogeography, 33:1578-1584 [featured article]

McClain, C.R. and J. Crouse (2006) The influence of ecological role on bathymetric patterns of deep-sea species: size clines in parasitic gastropods. Marine Ecology Progress Series, 320:161-167

Rex, M.A., R.J. Etter, J.S. Morris, J. Crouse, C.R. McClain, N.A. Johnson, C.T. Stuart, R. Thies, R. Avery (2006) Global bathymetric patterns of standing stock and body size in the deep-sea benthos. Marine Ecology Progress Series, 317:1-8 [featured article]


McClain, C.R., E.P. White, and A.H. Hurlbert (2007) Challenges in the application of geometric constraint models. Global Ecology and Biogeography, 16:257-264

Johnson, N.A., M.A. Rex, J.W. Campbell, C.R. McClain, T.S. Moore, and M.D. Dowell (2007) Using SeaWiFS satellite imagery to predict standing stock of the deep-sea macrobenthos. Deep-Sea Research, 54:1350-1360

McClain, C.R. (2007). Guest Editorial: Seamounts: Identity Crisis or Split Personality? Journal of Biogeography, 34:2001-2008


McClain, C.R. and J.C. Nekola (2008) The role of local-scale process on terrestrial and deep-sea gastropod body size distributions across multiple scales. Evolutionary Ecology Research, 10:129-146


Payne, J.L., A.G. Boyer, J.H. Brown, S. Finnegan, M. Kowalewski, R.A. Krause Jr., S.K. Lyons, C.R. McClain, D.W. McShea, P.M. Novack-Gottshall, F.A. Smith, J.A. Stempien, and S.C. Wang (2009) Two-phase increase in the maximum size of life over 3.5 billion years relfects biological innovation and environmental oppurtunity. Proceedings of the National Academy of Science, U.S.A., 1:24-27

McClain, C.R., L. Lundsten, M. Ream, J. Barry, and A. DeVogelaere (2009). Endemicity, Biogeography, Composition, and Community Structure On a Northeast Pacific Seamount. PLoS One, 4:e4141

McClain, C.R. and A.G. Boyer (2009) Biodiversity and body size are linked across metazoans. Proceedings of Royal Society B: Biological Sciences, 276: 2209-2215 [Highlighted by Faculty of 1000]

Lundsten, L., C.R. McClain, J.P. Barry, G.M. Caillet, D.A. Clague, A. DeVogelaere, and J.B. Geller (2009) Ichthyofauna on three seamounts off southern and central California. Marine Ecology Progress Series, 398:223-232

McClain, C.R., M.A. Rex, and R.J. Etter. (2009) Macroecological Patterns in the Deep Sea. In Marine Marcoecology. Eds. K. Roy and J. Whitman. University of Chicago Press


McClain, C.R. and Barry, J.P (2010) Habitat heterogeneity, biogenic disturbance, and resource availability work in concert to regulate biodiversity in deep submarine canyons. Ecology, 91:964-976

McClain, C.R. and S. Hardy (2010) The dynamcis of biogeographic ranges in the deep sea. Proceeding of the Royal Society, B, 282:3533-3546

McClain, C.R., L. Lundsten, J. Barry, and A. DeVogelaere (2010) Assemblage structure, but not diversity or density, change with depth on a northeast Pacific seamount. Marine Ecology, 31:1-12

Rex, M.A., C.T. Stuart, R.J. Etter, and C.R. McClain (2010) Biogeography of the deep-sea gastropod Oocorys sulcata Fischer 1884. Journal of Conchology 40: 287-290


Payne, J.L., C.R. McClain, A.G. Boyer, J.H. Brown, S. Finnegan, M. Kowalewski, R.A. Krause Jr., S.K. Lyons, D.W. McShea, P.M. Novack-Gottshall, F.A. Smith, J.A. Stempien, and S.C. Wang (2011) The evolutionary consequences of oxygenic photosynthesis: a body size perspective. Photosynthesis Research, 107:37-57

Finnegan, S., C.R. McClain, M.A. Kosnik, and J.L. Payne (2011) Escargot through time: an energetic comparison of marine gastropod assemblages before and after the Mesozoic Marine Revolution. Paleobiology, 37:252-269

McClain, C.R., J.Nekola, L.Kuhnz, and J. Barry (2011) Local-scale faunal turnover on the deep Pacific floor. Marine Ecology Progress Series, 442:193-200

Tittensor, D.P., M.A. Rex, C.T. Stuart, C.R. McClain, and C.R. Smith (2011) Species-energy relationships in deep-sea mollusks, Biology Letters, 7:718–722.

Kowalewski, M., J.L. Payne, F.A. Smith, S.C. Wang, D.W. McShea, S. Xiao, P.M. Novack-Gottshall, C.R. McClain, R.A. Krause, A.G. Boyer, S. Finnegan, S.K. Lyons, J.A. Stempien, J. Alroy, and P.A. Spaeth (2011) The Geozoic Supereon. Paleos 26:251-255


McClain, C.R., J.C. Stegen, and A.H. Hurlbert (2012) Dispersal, Niche Dynamics, and Oceanic Patterns in Beta-Diversity in Deep-Sea Bivalves. Proceeding of the Royal Society, B,  22:1993-2002

McClain, C.R., P. Unmack, J. Jackson-Ricketts, and T. Gullet (2012). Increased Energy Promotes Size-Based Niche Availability in Marine Mollusks. Evolution,  66:2204-2215

Thaler, A.D., K.A. Zelnio, A. Freitag, R. MacPherson, D. Shiffman, H. Bik, M.C. Goldstein, C. McClain (2012). Digital Environmentalism: Tools and strategies for the evolving online ecosystem. In D. Gallagher (ed.) Environmental Leadership: A Reference Handbook. SAGE Reference (Preprint pdf).

McClain, C.R., A.P. Allen, D.P. Tittensor, and M.A. Rex (2012) The Energetics of Life on the Deep Seafloor. Proceedings of the National Academy of Science, U.S.A. 109: 15366-15371

Harnik, P.G., H.K. Lotze, S.C. Anderson, Z.V. Finkel, S. Finnegan, D.R. Lindberg, L.H. Liow, R. Lockwood, C.R. McClain, J.L. McGuire, A. O’Dea, J.M. Pandolfi, C. Simpson, and D.P. Tittensor (2012) Extinctions in ancient and modern seas. Trends in Ecology and Evolution. 27:608-617

Payne, J.L., F.A. Smith, M. Kowalewski, R.A. Krause, A.G. Boyer, C.R. McClain, S. Finnegan, P.M. Novack-Gottshall, and L. Sheble (2012) A lack of attribution: closing the citation gap through a reform of citation and indexing practices. Taxon


Rodrigo A., S. Alberts, K. Cranston, J. Kingsolver, H. Lapp, C. McClain, R. Smith, T. Vision, J. Weintraub, and B. Weigmann(2013) Science Incubators: Synthesis Centers and Their Role in the Research Ecosystem. PLoS Biology 11(1): e1001468

McClain C.R., P.A.P. Durst, A.G. Boyer, C.D. Francis (2013) Unravelling the determinants of insular body size shifts. Biology Letters 9: 20120989

Brault, S., C.T. Stuart, M.C. Wagstaff, C.R. McClain, J.A. Allen, M.A. Rex (2013) Contrasting patterns of α- and β-diversity in deep-sea bivalves of the eastern and western North Atlantic. Deep-Sea Research II 92:157-164


McClain, C.R., R. Filler, C. Botero, and J.R. Auld (2014) Energy availability predicts diversity in gastropod reproductive strategies. Proceedings of the Royal Society, B 281:20140400

McClain, C.R. and J. Barry (2014) Beta-Diversity on Deep-Sea Wood Falls Reflects Gradients in Energy Availability. Biology Letters 10:2014012

Payne, J., N. Heim, M. Knope, C.R. McClain (2014) Metabolic dominance of bivalves predates brachiopod diversity decline by 200 million years. Proceedings of the Royal Society, B 281:20133122


Nunn, C.L., S.C. Alberts, C.R. McClain, S. Meshnick, T.J. Vison, B.M. Weigmann, A.G. Rodrigo (2015) Linking Evolution, Ecology, and Health: TriCEM. Bioscience online early

McClain, C.R. and M.A. Rex (2015) Toward a Conceptual Understanding of β-Diversity in the Deep-Sea Benthos. Annual Review of Ecology and Systematics in press

Orzechowski, E., R. Lockwood, J. Byrnes, S. Anderson, S. Finnegan, P. Harnik, Z.V. Finkel, D. Lindberg, L. Liow, H. Lotze, C.R. McClain, J. McGuire, A. O’Dea, J. Pandolfi, C. Simpson, and D. Tittensor (2015) Determinants of extinction selectivity over the last 500 million years: A meta-analysis of marine bivalves and gastropods. Global Change Biology in press

Finnegan, S., S.C. Anderson, P.G. Harnik, C. Simpson, D.P. Tittensor, J.E. Byrnes, Z.V. Finkel, D.R. Lindberg, L.H. Liow, R. Lockwood, H. Lotze, C.R. McClain, J.L. McGurie, A. O’Dean, and J.M. Pandolfi (2015) Paleontological baselines for evaluating extinction risk in the modern oceans. Science 348:567-570

McClain, C.R. and T. Schlacher (2015) On some hypotheses of diversity of animal life at great depths on the seafloor. Marine Ecology online early

Bik, H.M., A.D.M. Dove, M.C. Goldstein, R. Helm, R. MacPherson, K. Martini, A. Warneke, C.R. McClain (2015) Ten simple rules for effective online outreach. PLOS Computational Biology 11:e1003906

McClain, C.R., M.A. Balk, M.C. Benfield, T.A. Branch, C. Chen, J. Cosgrove, A. Dove, R.R. Helm, F.G.E. Hochberg, L.C. Gaskins, F.B. Lee, S.E. McMurray, C. Schanche, S.N. Stone, and A.D. Thaler (2015) Sizing ocean giants: patterns of intraspecific size variation in marine megafauna. Peer J, 2:e715

Martin, L.J., R.R. Dunn, R. Adams, A. Bateman, H.M. Bik, J. Hawks, S. Hird, D. Hughes, S. Kembel, K. Kinney, S.-O. Kolokotronis, C. Levy, C.R. McClain, J.F. Meadow, R. Medina, G. Mhuireach, C. Moreau, J. Munshi-South, C. Palmer, L. Popova, C. Schal, M. Taubel, M. Trautwein, J. Ugalde, (2015) Evolution of the indoor biome Trends in Ecology and Evolution 30:223-232

McClain C.R. and L. Lundsten (2015) Assemblage structure is related to slope and depth on a deep offshore Pacific seamount chain. Marine Ecology 36:210-220

Popular Writings

McClain, C.R. (2005) The Gold-Footed Snail. American Conchologist, 32:32-35

McClain, C.R. and J.C. Nekola (2007) Notes on two cases of molluscan biogeography. American Conchologist, 35:4-7

McClain, C.R. and P. Etnoyer (2007) Our Ocean Futures: The Glass Half-Empty and The Glass Half-Full. In The Open Laboratory: The Best Science Writing on Blogs 2007. Eds. R.A. Cartwright and B. Zivkovic.

McClain, C.R. (2008) Giants and dwarfs in the deep sea. Cosmos, 21

McClain, C.R. (2010) Archangel with Aqualung A review of Matsen’s The Sea King. American Scientist

McClain, C.R. (2010) An Empire Lacking Food. American Scientist

McClain, C.R. (2011) The Extinction of Species and Scientists Studying Them, Wired

McClain, C.R. (2011) Scientists Take Charles Darwin on the Road, Miller-McCune

McClain, C.R. (2012) What the deep seas tell us about life on other planets io9

McClain, C.R. (2012) On Sticking Your Arm Into an Underwater Cavern and Hoping a Catfish Bites You Mental Floss

McClain, C.R. (2013) Where There’s Heat, There Are Cockroaches Scientific American

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